==== Secondary Structure Definition by the program DSSP, updated CMBI version by ElmK / April 1,2000 ==== DATE=20-JUL-2011 . REFERENCE W. KABSCH AND C.SANDER, BIOPOLYMERS 22 (1983) 2577-2637 . HEADER HISTOCOMPATIBILITY ANTIGEN 15-FEB-94 1DLH . COMPND 2 MOLECULE: CLASS II HISTOCOMPATIBILITY ANTIGEN (HLA-DR1) (AL . SOURCE 2 ORGANISM_SCIENTIFIC: HOMO SAPIENS; . AUTHOR L.J.STERN . 761 7 6 6 0 TOTAL NUMBER OF RESIDUES, NUMBER OF CHAINS, NUMBER OF SS-BRIDGES(TOTAL,INTRACHAIN,INTERCHAIN) . 33880.0 ACCESSIBLE SURFACE OF PROTEIN (ANGSTROM**2) . 555 72.9 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(J) , SAME NUMBER PER 100 RESIDUES . 4 0.5 TOTAL NUMBER OF HYDROGEN BONDS IN PARALLEL BRIDGES, SAME NUMBER PER 100 RESIDUES . 287 37.7 TOTAL NUMBER OF HYDROGEN BONDS IN ANTIPARALLEL BRIDGES, SAME NUMBER PER 100 RESIDUES . 8 1.1 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(I-5), SAME NUMBER PER 100 RESIDUES . 4 0.5 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(I-4), SAME NUMBER PER 100 RESIDUES . 10 1.3 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(I-3), SAME NUMBER PER 100 RESIDUES . 0 0.0 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(I-2), SAME NUMBER PER 100 RESIDUES . 0 0.0 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(I-1), SAME NUMBER PER 100 RESIDUES . 0 0.0 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(I+0), SAME NUMBER PER 100 RESIDUES . 0 0.0 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(I+1), SAME NUMBER PER 100 RESIDUES . 94 12.4 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(I+2), SAME NUMBER PER 100 RESIDUES . 64 8.4 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(I+3), SAME NUMBER PER 100 RESIDUES . 92 12.1 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(I+4), SAME NUMBER PER 100 RESIDUES . 16 2.1 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(I+5), SAME NUMBER PER 100 RESIDUES . 1 2 3 4 5 6 7 8 9 10 11 12 13 14 15 16 17 18 19 20 21 22 23 24 25 26 27 28 29 30 *** HISTOGRAMS OF *** . 0 0 0 2 1 1 0 1 2 0 0 0 1 0 0 0 0 0 0 0 0 2 0 0 0 0 0 0 0 0 RESIDUES PER ALPHA HELIX . 2 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 PARALLEL BRIDGES PER LADDER . 8 8 8 5 1 10 2 4 3 0 2 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 ANTIPARALLEL BRIDGES PER LADDER . 0 0 3 5 0 0 1 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 LADDERS PER SHEET . # RESIDUE AA STRUCTURE BP1 BP2 ACC N-H-->O O-->H-N N-H-->O O-->H-N TCO KAPPA ALPHA PHI PSI X-CA Y-CA Z-CA 1 3 A E 0 0 127 0, 0.0 196,-0.2 0, 0.0 3,-0.1 0.000 360.0 360.0 360.0-162.1 19.2 65.4 8.8 2 4 A E - 0 0 96 194,-2.1 2,-0.3 1,-0.5 195,-0.2 0.685 360.0 -0.1-128.5 -38.1 21.6 67.1 11.1 3 5 A H E -A 196 0A 31 193,-1.1 193,-2.9 196,-0.1 -1,-0.5 -0.980 50.4-155.5-155.6 154.7 19.9 68.5 14.2 4 6 A V E -A 195 0A 18 -2,-0.3 21,-2.6 191,-0.2 2,-0.4 -0.995 1.4-163.7-135.0 138.2 16.5 68.8 15.7 5 7 A I E -AB 194 24A 9 189,-2.3 189,-2.3 -2,-0.4 2,-0.4 -0.959 17.8-168.9-117.9 135.8 15.3 71.3 18.2 6 8 A I E -AB 193 23A 15 17,-2.0 17,-1.9 -2,-0.4 2,-0.7 -0.999 21.2-159.2-134.3 136.9 12.1 70.3 20.0 7 9 A Q E +AB 192 22A 0 185,-2.0 185,-1.6 -2,-0.4 2,-0.5 -0.951 26.6 178.4-110.7 100.8 9.6 72.3 22.2 8 10 A A E +AB 191 21A 5 13,-1.9 13,-2.3 -2,-0.7 2,-0.3 -0.950 7.0 162.0-113.2 135.6 7.9 69.6 24.1 9 11 A E E +AB 190 20A 0 181,-1.7 181,-2.8 -2,-0.5 2,-0.3 -0.960 6.5 172.3-144.9 156.6 5.4 70.4 26.7 10 12 A F E -AB 189 19A 6 9,-2.0 9,-1.9 -2,-0.3 2,-0.4 -0.981 20.5-146.0-159.7 162.4 2.6 68.3 28.3 11 13 A Y E -AB 188 18A 20 177,-2.4 177,-1.9 -2,-0.3 2,-0.4 -0.999 15.9-159.0-136.4 140.2 0.0 68.3 31.0 12 14 A L E >> -AB 187 17A 0 5,-2.6 5,-1.5 -2,-0.4 4,-1.0 -0.957 13.3-128.5-126.1 138.2 -1.0 65.3 33.1 13 15 A N E 45S+A 186 0A 19 173,-2.1 173,-2.4 -2,-0.4 2,-0.0 -0.947 82.7 21.1-125.2 153.0 -4.2 64.7 35.1 14 16 A P T 45S+ 0 0 43 0, 0.0 -1,-0.1 0, 0.0 171,-0.1 -0.916 127.3 55.2 -63.4 -30.7 -5.3 63.8 37.8 15 17 A D T 45S- 0 0 58 1,-0.2 -2,-0.3 2,-0.1 3,-0.1 0.646 92.2-144.9 -36.5 -30.7 -1.8 64.9 38.9 16 18 A Q T <5 + 0 0 127 -4,-1.0 2,-0.4 1,-0.2 -3,-0.2 0.839 48.1 145.1 64.1 31.4 -2.4 68.3 37.3 17 19 A S E < +B 12 0A 45 -5,-1.5 -5,-2.6 48,-0.0 2,-0.3 -0.864 24.7 179.1-109.1 136.1 1.3 68.5 36.3 18 20 A G E -B 11 0A 17 -2,-0.4 2,-0.3 -7,-0.3 -7,-0.2 -0.935 9.7-160.7-133.3 157.6 2.8 70.1 33.2 19 21 A E E -B 10 0A 18 -9,-1.9 -9,-2.0 -2,-0.3 2,-0.4 -0.976 3.2-168.8-137.7 149.7 6.2 70.5 31.8 20 22 A F E +B 9 0A 1 12,-0.3 12,-1.7 -2,-0.3 2,-0.3 -0.993 25.9 145.9-140.3 124.8 7.6 72.9 29.1 21 23 A M E -BC 8 31A 0 -13,-2.3 -13,-1.9 -2,-0.4 2,-0.4 -0.979 38.3-130.8-156.9 166.0 11.1 72.5 27.6 22 24 A F E -BC 7 30A 0 8,-2.2 7,-3.1 -2,-0.3 8,-0.8 -0.930 21.7-168.4-121.7 148.3 13.4 72.7 24.7 23 25 A D E -BC 6 28A 5 -17,-1.9 -17,-2.0 -2,-0.4 2,-0.5 -0.983 10.4-161.1-137.8 138.4 15.7 70.0 23.5 24 26 A F E > S-BC 5 27A 1 3,-2.9 3,-2.3 -2,-0.4 -19,-0.2 -0.985 80.1 -26.0-123.2 119.3 18.5 70.1 21.0 25 27 A D T 3 S- 0 0 53 -21,-2.6 -1,-0.1 -2,-0.5 -20,-0.1 0.887 126.5 -47.9 45.7 51.2 19.4 66.7 19.6 26 28 A G T 3 S+ 0 0 16 -22,-0.2 2,-0.4 1,-0.1 -1,-0.3 0.250 117.6 105.2 82.1 -17.7 18.2 64.8 22.6 27 29 A D E < S-C 24 0A 2 -3,-2.3 -3,-2.9 304,-0.1 -1,-0.1 -0.878 76.5-112.4 -99.2 137.0 19.9 67.0 25.3 28 30 A E E -C 23 0A 7 -2,-0.4 -5,-0.3 302,-0.3 14,-0.1 -0.376 24.9-171.5 -67.8 134.5 17.7 69.5 27.2 29 31 A I E - 0 0 0 -7,-3.1 13,-1.9 1,-0.5 14,-0.4 0.890 64.6 -45.5 -89.9 -48.8 18.4 73.2 26.5 30 32 A F E -CD 22 41A 0 -8,-0.8 -8,-2.2 11,-0.3 -1,-0.5 -0.925 50.4-152.5-168.6 170.2 16.2 74.6 29.3 31 33 A H E -CD 21 40A 42 9,-1.2 9,-3.8 -2,-0.3 2,-0.5 -0.964 26.5-108.1-152.4 155.8 12.7 74.2 30.8 32 34 A V E - D 0 39A 3 -12,-1.7 2,-0.9 -2,-0.3 -12,-0.3 -0.750 23.5-129.6 -94.3 122.1 10.4 76.6 32.7 33 35 A D E >> - D 0 38A 43 5,-2.9 4,-1.9 -2,-0.5 5,-0.8 -0.601 16.3-160.9 -71.6 109.6 9.8 76.2 36.4 34 36 A M T 45S+ 0 0 46 -2,-0.9 -1,-0.2 -16,-0.2 -15,-0.1 0.859 88.1 43.9 -56.5 -41.1 6.0 76.2 36.6 35 37 A A T 45S+ 0 0 75 1,-0.2 -1,-0.2 -3,-0.1 -2,-0.1 0.989 123.3 30.7 -69.5 -67.5 6.3 77.0 40.3 36 38 A K T 45S- 0 0 144 2,-0.1 -2,-0.2 1,-0.0 -1,-0.2 0.639 101.5-134.5 -68.3 -15.1 9.0 79.7 40.4 37 39 A K T <5 + 0 0 120 -4,-1.9 2,-0.3 1,-0.2 -3,-0.2 0.977 62.2 121.8 56.9 60.3 7.8 80.7 37.0 38 40 A E E < -D 33 0A 78 -5,-0.8 -5,-2.9 16,-0.1 2,-0.3 -0.981 66.5-109.5-153.5 137.8 11.4 81.0 35.7 39 41 A T E -D 32 0A 28 -2,-0.3 2,-0.7 13,-0.3 -7,-0.3 -0.466 27.4-152.9 -74.1 133.8 13.3 79.4 33.0 40 42 A V E -D 31 0A 46 -9,-3.8 -9,-1.2 -2,-0.3 -1,-0.1 -0.873 6.2-147.3-111.5 104.9 15.9 77.0 34.1 41 43 A W E -D 30 0A 43 -2,-0.7 -11,-0.3 -11,-0.2 3,-0.2 -0.330 12.5-136.1 -64.7 150.4 18.9 76.5 31.7 42 44 A R S S+ 0 0 49 -13,-1.9 2,-0.4 1,-0.3 -12,-0.1 0.752 98.4 25.1 -80.3 -34.4 20.5 73.0 31.7 43 45 A L S > S- 0 0 28 -14,-0.4 3,-2.9 1,-0.1 4,-0.4 -0.987 85.2-134.5-128.2 117.3 24.1 74.2 31.8 44 46 A E G >> S+ 0 0 172 -2,-0.4 3,-1.8 1,-0.3 4,-0.7 0.782 103.3 73.0 -39.8 -37.8 24.2 77.8 33.4 45 47 A E G >4 S+ 0 0 87 1,-0.3 3,-1.0 2,-0.2 -1,-0.3 0.815 86.7 64.3 -45.8 -41.1 26.6 78.7 30.5 46 48 A F G X> S+ 0 0 2 -3,-2.9 4,-1.7 1,-0.2 3,-0.7 0.784 96.1 56.3 -56.3 -32.7 23.5 78.6 28.3 47 49 A G G <4 S+ 0 0 31 -3,-1.8 -1,-0.2 -4,-0.4 -2,-0.2 0.707 91.6 69.7 -75.4 -24.1 21.9 81.5 30.2 48 50 A R G << S+ 0 0 170 -3,-1.0 -1,-0.2 -4,-0.7 -2,-0.1 0.125 114.4 26.2 -79.3 24.6 24.8 83.9 29.6 49 51 A F T <4 S+ 0 0 97 -3,-0.7 2,-0.2 1,-0.5 -2,-0.2 0.520 120.1 34.7-150.4 -51.0 23.9 84.0 25.9 50 52 A A < - 0 0 5 -4,-1.7 -1,-0.5 215,-0.1 2,-0.3 -0.747 59.2-157.8-117.9 164.3 20.2 83.4 25.2 51 53 A S B -i 372 0B 52 320,-2.1 322,-2.8 -2,-0.2 2,-0.3 -0.869 5.6-166.7-137.9 168.8 17.0 84.1 27.0 52 54 A F - 0 0 17 -2,-0.3 2,-1.2 320,-0.2 -13,-0.3 -0.941 27.9-121.7-153.3 141.6 13.4 82.8 27.2 53 55 A E >> - 0 0 110 320,-0.4 3,-3.5 -2,-0.3 4,-0.6 -0.691 18.4-156.5 -87.5 94.5 10.3 84.4 28.8 54 56 A A H 3> S+ 0 0 14 -2,-1.2 4,-1.5 1,-0.3 -1,-0.2 0.691 89.2 65.4 -44.5 -21.2 9.1 81.9 31.3 55 57 A Q H 3> S+ 0 0 136 1,-0.2 4,-1.7 2,-0.2 -1,-0.3 0.795 87.6 68.7 -73.7 -27.7 5.6 83.4 31.1 56 58 A G H <> S+ 0 0 14 -3,-3.5 4,-2.1 1,-0.2 3,-0.4 0.971 103.2 44.6 -53.9 -47.1 5.4 82.2 27.5 57 59 A A H X S+ 0 0 0 -4,-0.6 4,-2.9 1,-0.2 5,-0.3 0.936 107.9 54.3 -64.1 -49.0 5.3 78.7 28.8 58 60 A L H X S+ 0 0 38 -4,-1.5 4,-0.9 1,-0.2 -1,-0.2 0.804 109.5 53.0 -56.0 -27.0 2.8 79.2 31.7 59 61 A A H >X S+ 0 0 50 -4,-1.7 3,-1.4 -3,-0.4 4,-1.0 0.997 109.8 45.3 -66.8 -64.0 0.6 80.7 29.0 60 62 A N H 3X S+ 0 0 5 -4,-2.1 4,-2.0 1,-0.3 -2,-0.2 0.771 108.1 57.4 -48.3 -40.1 0.9 77.6 26.7 61 63 A I H 3X S+ 0 0 11 -4,-2.9 4,-3.0 2,-0.2 -1,-0.3 0.862 96.3 64.0 -66.1 -31.6 0.3 75.1 29.5 62 64 A A H X S+ 0 0 32 -4,-2.3 4,-1.9 1,-0.2 3,-0.8 0.999 112.8 45.4 -65.6 -67.7 -11.4 68.2 26.0 71 73 A M H 3X S+ 0 0 0 -4,-2.9 4,-2.0 1,-0.3 6,-0.2 0.687 108.0 58.1 -46.9 -35.6 -9.5 65.3 24.5 72 74 A T H 3X>S+ 0 0 13 -4,-2.5 5,-2.3 2,-0.2 4,-0.9 0.922 108.0 47.8 -62.8 -49.8 -10.2 63.0 27.5 73 75 A K H X<5S+ 0 0 119 -4,-1.3 3,-2.5 -3,-0.8 -2,-0.2 0.998 112.4 46.1 -58.8 -56.4 -13.8 63.5 27.0 74 76 A R H 3<5S+ 0 0 63 -4,-1.9 -1,-0.2 1,-0.3 -2,-0.2 0.848 104.5 61.7 -59.0 -27.9 -13.8 62.8 23.4 75 77 A S H 3<5S- 0 0 13 -4,-2.0 -1,-0.3 -5,-0.2 -2,-0.2 0.763 117.7-115.5 -63.6 -24.9 -11.6 59.7 23.8 76 78 A N T <<5 - 0 0 139 -3,-2.5 -3,-0.2 -4,-0.9 -2,-0.2 0.642 64.2 -63.0 91.0 24.1 -14.4 58.4 25.9 77 79 A Y S -O 99 0E 113 3,-0.6 3,-1.3 -2,-0.4 82,-0.0 -0.913 40.9 -94.3-118.5 145.4 39.6 54.5 39.7 97 99 A L T 3 S+ 0 0 58 -2,-0.4 57,-0.1 1,-0.2 -1,-0.0 -0.290 107.9 0.9 -61.8 143.8 39.8 54.3 43.5 98 100 A R T 3 S+ 0 0 197 55,-1.0 -1,-0.2 1,-0.2 56,-0.1 0.450 101.8 119.6 55.9 0.8 39.8 57.7 45.2 99 101 A E B < S-O 96 0E 97 -3,-1.3 -3,-0.6 1,-0.0 -1,-0.2 -0.816 73.2-115.9 -97.6 102.8 39.4 59.3 41.8 100 102 A P + 0 0 100 0, 0.0 52,-0.2 0, 0.0 2,-0.2 0.093 47.1 158.8 -38.7 138.1 36.0 61.3 41.9 101 103 A N E - L 0 151D 12 50,-1.8 50,-2.3 48,-0.1 2,-0.3 -0.705 32.5-120.2-146.9-172.2 33.3 60.1 39.5 102 104 A V E - L 0 150D 33 48,-0.3 -10,-2.0 -2,-0.2 2,-0.5 -0.985 12.3-139.9-143.1 143.2 29.5 60.3 39.1 103 105 A L E -KL 91 149D 0 46,-2.3 46,-1.9 -2,-0.3 2,-0.5 -0.949 18.1-155.7-105.8 120.2 26.9 57.5 38.9 104 106 A I E -KL 90 148D 0 -14,-0.8 -14,-2.7 -2,-0.5 2,-0.8 -0.844 4.2-156.0 -98.1 120.8 24.1 58.1 36.3 105 107 A a E -KL 89 147D 0 42,-2.7 42,-1.0 -2,-0.5 2,-0.6 -0.841 15.8-158.1 -96.3 107.8 20.7 56.3 36.9 106 108 A F E -KL 88 146D 43 -18,-2.6 -18,-2.2 -2,-0.8 2,-0.5 -0.815 10.0-172.4 -94.7 120.0 19.2 56.1 33.4 107 109 A I E +KL 87 145D 0 38,-2.7 38,-2.4 -2,-0.6 -20,-0.2 -0.952 18.2 168.1-110.9 117.0 15.4 55.8 33.3 108 110 A D E +K 86 0D 16 -22,-1.0 -22,-1.3 -2,-0.5 36,-0.2 -0.671 41.4 22.9-125.6 175.2 14.0 55.1 29.7 109 111 A K E S+ 0 0 45 -2,-0.2 2,-0.2 -24,-0.2 35,-0.1 0.570 80.9 134.6 43.2 33.0 10.9 54.0 27.8 110 112 A F E - L 0 143D 0 33,-1.0 33,-2.7 -3,-0.1 -1,-0.1 -0.687 37.3-146.2-114.7 160.2 8.2 54.9 30.3 111 113 A T B S+J 83 0C 0 -28,-1.8 -28,-3.1 -2,-0.2 56,-0.1 -0.903 71.0 47.7-163.2 141.0 4.8 56.6 30.5 112 114 A P S S- 0 0 0 0, 0.0 2,-3.1 0, 0.0 -1,-0.2 0.487 86.8-120.9 -66.1 150.9 3.0 58.5 32.4 113 115 A P S S+ 0 0 3 0, 0.0 2,-0.3 0, 0.0 22,-0.1 -0.296 79.4 107.5 -66.9 64.0 5.8 60.9 33.2 114 116 A V + 0 0 35 -2,-3.1 52,-0.3 2,-0.0 2,-0.3 -0.989 36.5 144.0-139.6 136.3 5.7 60.3 37.0 115 117 A V - 0 0 19 -2,-0.3 2,-0.6 50,-0.1 50,-0.2 -0.928 38.4-136.1-174.3 141.0 8.4 58.3 38.8 116 118 A N E -P 164 0F 97 48,-2.4 48,-2.2 -2,-0.3 2,-0.5 -0.930 22.1-173.4-116.9 116.3 10.2 58.3 42.2 117 119 A V E +P 163 0F 30 -2,-0.6 2,-0.3 46,-0.2 46,-0.2 -0.916 4.5 178.0-114.7 131.1 14.0 57.8 42.1 118 120 A T E -P 162 0F 40 44,-1.9 44,-3.2 -2,-0.5 2,-0.4 -0.953 20.2-142.3-127.7 138.8 16.1 57.4 45.3 119 121 A W E -P 161 0F 8 7,-0.5 7,-1.4 -2,-0.3 2,-0.4 -0.816 19.4-168.2 -98.6 151.5 19.8 56.8 45.5 120 122 A L E -PQ 160 125F 18 40,-2.2 40,-2.0 -2,-0.4 2,-0.5 -0.998 11.5-167.4-141.8 131.7 20.9 54.5 48.2 121 123 A R E > S-PQ 159 124F 73 3,-3.2 3,-1.1 -2,-0.4 38,-0.2 -0.985 84.5 -9.7-119.6 117.3 24.5 54.0 49.3 122 124 A N T 3 S- 0 0 72 36,-2.9 -1,-0.2 -2,-0.5 37,-0.1 0.894 132.2 -52.4 61.0 43.1 24.8 50.9 51.5 123 125 A G T 3 S+ 0 0 39 35,-0.3 -1,-0.2 1,-0.2 581,-0.1 0.174 119.5 106.0 81.7 -15.8 21.1 50.4 51.8 124 126 A K E < S-Q 121 0F 146 -3,-1.1 -3,-3.2 1,-0.1 -1,-0.2 -0.775 75.6-104.5-106.0 142.1 20.6 54.1 53.0 125 127 A P E -Q 120 0F 93 0, 0.0 2,-0.4 0, 0.0 -5,-0.2 -0.141 34.1-158.7 -58.6 150.7 19.2 57.1 51.1 126 128 A V + 0 0 13 -7,-1.4 -7,-0.5 1,-0.1 0, 0.0 -0.991 28.2 165.6-141.0 129.7 21.6 59.7 49.7 127 129 A T > + 0 0 118 -2,-0.4 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