==== Secondary Structure Definition by the program DSSP, updated CMBI version by ElmK / April 1,2000 ==== DATE=20-JUL-2011 . REFERENCE W. KABSCH AND C.SANDER, BIOPOLYMERS 22 (1983) 2577-2637 . HEADER LYASE 27-APR-07 2PP0 . COMPND 2 MOLECULE: L-TALARATE/GALACTARATE DEHYDRATASE; . SOURCE 2 ORGANISM_SCIENTIFIC: SALMONELLA TYPHIMURIUM; . AUTHOR A.A.FEDOROV,E.V.FEDOROV,W.S.YEW,J.A.GERLT,S.C.ALMO . 1185 3 0 0 0 TOTAL NUMBER OF RESIDUES, NUMBER OF CHAINS, NUMBER OF SS-BRIDGES(TOTAL,INTRACHAIN,INTERCHAIN) . 44289.0 ACCESSIBLE SURFACE OF PROTEIN (ANGSTROM**2) . 786 66.3 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(J) , SAME NUMBER PER 100 RESIDUES . 77 6.5 TOTAL NUMBER OF HYDROGEN BONDS IN PARALLEL BRIDGES, SAME NUMBER PER 100 RESIDUES . 134 11.3 TOTAL NUMBER OF HYDROGEN BONDS IN ANTIPARALLEL BRIDGES, SAME NUMBER PER 100 RESIDUES . 0 0.0 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(I-5), SAME NUMBER PER 100 RESIDUES . 3 0.3 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(I-4), SAME NUMBER PER 100 RESIDUES . 3 0.3 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(I-3), SAME NUMBER PER 100 RESIDUES . 0 0.0 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(I-2), SAME NUMBER PER 100 RESIDUES . 0 0.0 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(I-1), SAME NUMBER PER 100 RESIDUES . 0 0.0 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(I+0), SAME NUMBER PER 100 RESIDUES . 0 0.0 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(I+1), SAME NUMBER PER 100 RESIDUES . 40 3.4 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(I+2), SAME NUMBER PER 100 RESIDUES . 127 10.7 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(I+3), SAME NUMBER PER 100 RESIDUES . 334 28.2 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(I+4), SAME NUMBER PER 100 RESIDUES . 20 1.7 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(I+5), SAME NUMBER PER 100 RESIDUES . 1 2 3 4 5 6 7 8 9 10 11 12 13 14 15 16 17 18 19 20 21 22 23 24 25 26 27 28 29 30 *** HISTOGRAMS OF *** . 0 0 0 9 6 0 0 0 6 3 9 0 6 0 3 0 0 0 0 3 0 0 0 0 0 0 0 0 0 0 RESIDUES PER ALPHA HELIX . 5 13 2 4 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 PARALLEL BRIDGES PER LADDER . 5 0 6 0 0 0 3 2 1 0 0 3 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 ANTIPARALLEL BRIDGES PER LADDER . 0 2 3 0 0 0 2 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 LADDERS PER SHEET . # RESIDUE AA STRUCTURE BP1 BP2 ACC N-H-->O O-->H-N N-H-->O O-->H-N TCO KAPPA ALPHA PHI PSI X-CA Y-CA Z-CA 1 4 A S 0 0 148 0, 0.0 3,-0.1 0, 0.0 0, 0.0 0.000 360.0 360.0 360.0 141.3 25.9 41.7 86.1 2 5 A A - 0 0 110 1,-0.1 2,-0.3 0, 0.0 0, 0.0 0.884 360.0 -18.8 -64.7 -38.6 24.2 38.9 84.2 3 6 A N - 0 0 127 4,-0.1 2,-0.2 2,-0.0 -1,-0.1 -0.906 57.3-159.3-156.1-177.4 24.0 36.7 87.3 4 7 A S - 0 0 52 2,-0.4 4,-0.1 -2,-0.3 0, 0.0 -0.806 48.5 -77.8-152.7-168.9 25.3 36.1 90.9 5 8 A D S S+ 0 0 182 -2,-0.2 2,-0.0 2,-0.1 -2,-0.0 0.676 120.3 26.5 -76.8 -17.4 25.6 33.4 93.5 6 9 A A S S- 0 0 85 0, 0.0 -2,-0.4 0, 0.0 2,-0.3 0.182 103.1 -85.4-111.0-128.6 22.0 33.9 94.4 7 10 A V - 0 0 97 -4,-0.1 2,-0.3 -2,-0.0 -2,-0.1 -0.988 34.0-166.7-150.8 143.9 19.2 35.2 92.2 8 11 A T - 0 0 101 -2,-0.3 2,-0.4 -4,-0.1 -4,-0.0 -0.936 9.3-142.8-133.0 158.2 18.1 38.7 91.3 9 12 A Y - 0 0 186 -2,-0.3 2,-0.2 0, 0.0 -2,-0.0 -0.980 11.1-134.7-128.1 133.5 15.0 40.2 89.6 10 13 A A - 0 0 103 -2,-0.4 2,-0.4 1,-0.0 -2,-0.0 -0.554 21.9-139.5 -81.1 145.8 14.7 43.2 87.2 11 14 A K - 0 0 183 -2,-0.2 2,-0.7 2,-0.0 3,-0.0 -0.871 15.8-116.8-110.4 142.4 11.9 45.7 87.9 12 15 A A - 0 0 69 -2,-0.4 3,-0.2 1,-0.2 -2,-0.0 -0.669 14.5-164.4 -82.2 113.4 9.7 47.3 85.2 13 16 A A S S+ 0 0 117 -2,-0.7 2,-0.4 1,-0.3 -1,-0.2 0.744 91.2 38.4 -64.4 -22.3 10.1 51.1 85.1 14 17 A N + 0 0 114 1,-0.1 -1,-0.3 2,-0.1 0, 0.0 -0.916 61.9 175.0-134.3 106.6 6.9 51.0 83.0 15 18 A T + 0 0 106 -2,-0.4 2,-0.4 -3,-0.2 -1,-0.1 0.102 33.1 137.0 -96.8 20.8 4.2 48.5 84.1 16 19 A R - 0 0 168 1,-0.1 2,-0.2 3,-0.0 -2,-0.1 -0.572 46.0-144.9 -69.1 122.7 1.7 49.7 81.5 17 20 A T > - 0 0 61 -2,-0.4 4,-1.5 1,-0.1 3,-0.2 -0.594 23.6-126.7-100.0 156.4 0.1 46.5 80.1 18 21 A A H > S+ 0 0 81 1,-0.2 4,-1.6 -2,-0.2 -1,-0.1 0.850 111.4 59.9 -60.4 -34.7 -1.1 45.4 76.7 19 22 A A H 4 S+ 0 0 72 1,-0.2 -1,-0.2 2,-0.2 3,-0.0 0.914 108.2 40.7 -61.3 -45.9 -4.3 44.4 78.5 20 23 A E H 4 S+ 0 0 107 -3,-0.2 -1,-0.2 1,-0.2 -2,-0.2 0.705 121.0 41.4 -78.9 -20.4 -5.1 47.9 79.7 21 24 A T H < S+ 0 0 58 -4,-1.5 -2,-0.2 2,-0.0 -1,-0.2 0.582 85.7 116.2-104.7 -11.1 -4.1 49.8 76.6 22 25 A G S < S- 0 0 13 -4,-1.6 -3,-0.0 -5,-0.2 -5,-0.0 -0.134 70.4 -98.7 -61.8 154.1 -5.5 47.6 73.8 23 26 A D - 0 0 4 44,-0.1 2,-0.3 45,-0.0 80,-0.2 -0.071 36.8-144.3 -62.2 170.2 -8.2 48.7 71.4 24 27 A R - 0 0 65 77,-0.2 43,-3.0 78,-0.1 2,-0.3 -0.998 14.0-118.1-144.7 137.4 -11.8 47.7 72.0 25 28 A I E +A 66 0A 2 75,-2.2 74,-2.8 72,-0.4 41,-0.2 -0.597 36.1 166.8 -74.3 129.2 -14.7 46.7 69.7 26 29 A E E + 0 0 75 39,-2.5 367,-2.0 -2,-0.3 2,-0.3 0.558 57.5 30.2-119.8 -13.2 -17.6 49.2 70.1 27 30 A W E -AB 65 392A 2 38,-1.7 38,-2.4 365,-0.2 2,-0.3 -0.997 52.4-179.4-152.1 143.5 -19.9 48.6 67.2 28 31 A V E -AB 64 391A 0 363,-2.2 363,-2.8 -2,-0.3 2,-0.4 -0.994 4.2-170.7-144.5 135.0 -21.0 45.7 64.9 29 32 A K E -AB 63 390A 27 34,-2.4 34,-2.4 -2,-0.3 2,-0.4 -0.995 5.9-167.3-127.6 130.7 -23.4 45.6 62.0 30 33 A L E -AB 62 389A 0 359,-2.6 359,-2.0 -2,-0.4 2,-0.3 -0.934 5.9-173.1-118.0 143.5 -24.6 42.4 60.4 31 34 A S E -AB 61 388A 0 30,-2.2 30,-2.7 -2,-0.4 2,-0.6 -0.991 18.7-154.6-138.2 144.2 -26.5 42.1 57.1 32 35 A L E +AB 60 387A 0 355,-2.8 354,-2.9 -2,-0.3 355,-1.6 -0.972 29.0 171.6-115.6 112.9 -28.2 39.3 55.1 33 36 A A E -AB 59 385A 0 26,-3.2 26,-3.3 -2,-0.6 2,-0.5 -0.842 25.0-139.3-120.4 159.1 -28.2 40.2 51.5 34 37 A F E -A 58 0A 20 350,-1.9 24,-0.2 -2,-0.3 -2,-0.0 -0.983 6.2-153.6-124.3 123.8 -29.2 38.2 48.4 35 38 A L E -A 57 0A 4 22,-3.1 22,-2.1 -2,-0.5 2,-0.2 -0.854 20.6-140.8 -94.1 111.0 -27.3 38.3 45.1 36 39 A P E -A 56 0A 71 0, 0.0 2,-0.7 0, 0.0 20,-0.3 -0.509 3.3-147.4 -77.3 142.2 -29.6 37.5 42.2 37 40 A L E - 0 0 22 18,-2.8 4,-0.1 -2,-0.2 312,-0.0 -0.932 2.3-159.5-109.0 108.0 -28.3 35.4 39.4 38 41 A A E S+ 0 0 105 -2,-0.7 -1,-0.1 1,-0.2 18,-0.0 0.835 85.0 49.7 -54.7 -36.9 -30.0 36.5 36.1 39 42 A T E S- 0 0 90 -3,-0.1 -1,-0.2 15,-0.0 15,-0.1 -0.921 95.6-123.4-109.8 111.6 -29.1 33.1 34.5 40 43 A P E - 0 0 64 0, 0.0 2,-0.4 0, 0.0 15,-0.2 -0.279 30.8-136.3 -51.9 132.5 -30.1 30.1 36.6 41 44 A V E -A 54 0A 13 13,-2.6 13,-3.0 11,-0.1 2,-0.4 -0.822 22.4-175.8-104.1 131.8 -27.0 28.0 37.2 42 45 A S + 0 0 12 -2,-0.4 127,-1.3 11,-0.2 128,-0.8 -0.977 7.0 170.4-128.9 124.1 -26.8 24.2 36.9 43 46 A D >> - 0 0 14 -2,-0.4 4,-1.6 125,-0.2 3,-1.4 -0.408 61.8 -72.1-109.0-167.2 -23.8 21.9 37.6 44 47 A A H 3> S+ 0 0 0 151,-0.4 4,-1.8 1,-0.3 6,-0.3 0.779 127.8 68.2 -60.8 -27.4 -23.9 18.1 37.8 45 48 A K H 34>S+ 0 0 19 1,-0.2 6,-1.7 2,-0.2 5,-1.2 0.832 106.4 42.1 -60.5 -31.4 -25.8 18.2 41.1 46 49 A V H X45S+ 0 0 22 -3,-1.4 3,-1.6 4,-0.2 -2,-0.2 0.907 111.1 53.1 -80.9 -45.8 -28.7 19.6 39.2 47 50 A L H 3<5S+ 0 0 61 -4,-1.6 -2,-0.2 1,-0.3 -1,-0.2 0.721 108.5 51.1 -65.2 -20.5 -28.5 17.3 36.2 48 51 A T T 3<5S- 0 0 49 -4,-1.8 -1,-0.3 149,-0.1 -2,-0.2 0.455 118.8-111.7 -93.2 -2.6 -28.6 14.2 38.4 49 52 A G T < 5S+ 0 0 64 -3,-1.6 -3,-0.2 2,-0.2 -2,-0.1 0.463 85.8 117.8 89.0 0.6 -31.7 15.5 40.2 50 53 A R S -A 25 0A 0 4,-2.6 3,-1.9 -2,-0.4 -41,-0.2 -0.638 27.7-110.9-105.5 162.0 -13.2 49.9 67.2 67 70 A R T 3 S+ 0 0 99 -43,-3.0 -42,-0.1 1,-0.3 -1,-0.1 0.883 115.3 55.8 -52.2 -45.8 -12.8 52.6 69.9 68 71 A D T 3 S- 0 0 83 -44,-0.2 -1,-0.3 1,-0.1 -43,-0.1 0.259 125.4 -94.7 -79.1 14.9 -9.5 53.7 68.3 69 72 A G S < S+ 0 0 58 -3,-1.9 2,-0.2 1,-0.3 -2,-0.1 0.578 80.7 129.0 90.0 10.1 -11.0 54.4 64.9 70 73 A F - 0 0 62 2,-0.0 -4,-2.6 73,-0.0 2,-0.3 -0.555 31.5-174.2 -96.0 162.6 -10.4 51.1 63.0 71 74 A E E -C 65 0A 58 -6,-0.2 2,-0.3 -2,-0.2 -6,-0.2 -0.995 8.3-166.3-153.8 156.1 -12.9 49.0 61.1 72 75 A G E -C 64 0A 1 -8,-2.2 -8,-2.4 -2,-0.3 2,-0.3 -0.960 4.1-159.7-141.7 159.3 -13.2 45.8 59.3 73 76 A V E +C 63 0A 24 -2,-0.3 2,-0.3 62,-0.2 -10,-0.2 -0.993 19.4 156.3-140.8 142.5 -15.6 44.1 56.9 74 77 A G E -C 62 0A 1 -12,-1.8 -12,-2.5 -2,-0.3 2,-0.3 -0.952 14.0-164.9-155.9 170.3 -16.0 40.4 56.1 75 78 A F E -C 61 0A 10 -2,-0.3 2,-0.2 -14,-0.2 -14,-0.2 -0.993 12.5-163.5-160.0 162.8 -18.6 38.0 54.8 76 79 A S E -C 60 0A 0 -16,-1.8 -16,-1.9 -2,-0.3 2,-0.3 -0.706 21.6-158.8-132.8-174.7 -19.7 34.4 54.3 77 80 A Y E -C 59 0A 1 -2,-0.2 2,-0.3 -18,-0.2 -18,-0.2 -0.967 13.0-141.4-161.1 173.7 -22.3 33.1 52.0 78 81 A S E -C 58 0A 12 -20,-1.9 -20,-2.7 -2,-0.3 2,-0.7 -0.971 6.3-152.9-150.4 132.1 -24.7 30.3 51.1 79 82 A K S S- 0 0 34 -2,-0.3 2,-0.1 -22,-0.2 -22,-0.1 -0.944 81.0 -15.5-107.0 109.7 -25.8 28.8 47.8 80 83 A R S > S+ 0 0 72 -2,-0.7 3,-0.6 -24,-0.5 -22,-0.2 -0.429 128.1 4.0 85.4-165.2 -29.3 27.5 48.3 81 84 A A T 3 S+ 0 0 105 1,-0.2 -2,-0.2 -2,-0.1 0, 0.0 -0.168 103.5 70.8 -53.3 132.7 -30.6 27.0 51.9 82 85 A G T 3> + 0 0 18 -4,-0.1 4,-1.9 42,-0.0 -1,-0.2 0.119 64.6 91.6 150.4 -26.8 -28.4 28.2 54.7 83 86 A G H <> S+ 0 0 3 -3,-0.6 4,-2.3 1,-0.2 5,-0.2 0.885 87.2 52.2 -63.6 -40.3 -28.2 32.0 54.6 84 87 A Q H > S+ 0 0 105 1,-0.2 4,-2.4 2,-0.2 -1,-0.2 0.904 112.3 46.0 -63.1 -41.9 -31.1 32.5 57.0 85 88 A G H > S+ 0 0 30 2,-0.2 4,-2.0 1,-0.2 -1,-0.2 0.852 111.7 50.7 -70.2 -36.1 -29.5 30.2 59.6 86 89 A I H X S+ 0 0 0 -4,-1.9 4,-2.3 2,-0.2 -2,-0.2 0.933 114.6 43.4 -67.3 -46.4 -26.1 31.8 59.2 87 90 A Y H X S+ 0 0 10 -4,-2.3 4,-2.4 1,-0.2 5,-0.2 0.939 112.8 52.8 -63.8 -47.1 -27.4 35.3 59.7 88 91 A A H X S+ 0 0 54 -4,-2.4 4,-1.4 -5,-0.2 -1,-0.2 0.898 112.5 44.0 -57.0 -43.6 -29.7 34.3 62.5 89 92 A H H X S+ 0 0 27 -4,-2.0 4,-1.3 2,-0.2 3,-0.3 0.940 111.2 52.6 -69.3 -45.4 -26.9 32.7 64.5 90 93 A A H X S+ 0 0 0 -4,-2.3 4,-2.1 1,-0.3 3,-0.4 0.896 110.3 50.8 -55.1 -39.7 -24.4 35.5 63.9 91 94 A K H < S+ 0 0 109 -4,-2.4 4,-0.4 1,-0.2 -1,-0.3 0.839 107.3 52.9 -67.3 -33.7 -27.1 37.9 65.2 92 95 A E H < S+ 0 0 150 -4,-1.4 -1,-0.2 -3,-0.3 -2,-0.2 0.689 121.2 30.5 -78.1 -16.3 -27.6 35.8 68.3 93 96 A I H >< S+ 0 0 4 -4,-1.3 3,-1.8 -3,-0.4 -2,-0.2 0.581 91.5 92.5-115.8 -13.9 -23.9 35.8 69.2 94 97 A A G >< S+ 0 0 0 -4,-2.1 3,-2.6 1,-0.3 4,-0.4 0.812 73.8 68.3 -52.7 -37.5 -22.6 39.2 67.8 95 98 A D G > S+ 0 0 85 -4,-0.4 3,-1.0 1,-0.3 -1,-0.3 0.744 84.5 74.6 -58.4 -19.4 -23.1 41.1 71.0 96 99 A N G < S+ 0 0 41 -3,-1.8 -1,-0.3 1,-0.2 -2,-0.2 0.655 85.6 66.4 -66.0 -14.9 -20.2 39.0 72.5 97 100 A L G X S+ 0 0 0 -3,-2.6 3,-2.0 -4,-0.2 -72,-0.4 0.792 74.4 104.1 -77.2 -29.9 -17.9 41.1 70.4 98 101 A L T < S+ 0 0 61 -3,-1.0 -72,-0.2 -4,-0.4 3,-0.1 -0.304 89.8 12.2 -58.5 129.8 -18.5 44.4 72.3 99 102 A G T 3 S+ 0 0 44 -74,-2.8 -1,-0.3 1,-0.3 2,-0.2 0.387 96.5 127.4 86.4 -4.1 -15.6 45.3 74.6 100 103 A E S < S- 0 0 43 -3,-2.0 -75,-2.2 -75,-0.2 -1,-0.3 -0.565 70.2-110.3 -87.6 150.1 -13.2 42.7 73.2 101 104 A D > - 0 0 26 -77,-0.2 3,-1.6 -2,-0.2 -77,-0.2 -0.710 28.2-148.4 -76.8 113.0 -9.7 43.6 72.0 102 105 A P T 3 S+ 0 0 3 0, 0.0 -1,-0.2 0, 0.0 38,-0.1 0.637 89.5 68.0 -59.8 -14.5 -10.1 43.0 68.3 103 106 A N T 3 S+ 0 0 40 1,-0.2 2,-1.7 -80,-0.2 3,-0.3 0.730 81.4 75.3 -79.4 -21.8 -6.5 41.9 67.9 104 107 A D <> + 0 0 69 -3,-1.6 4,-2.4 1,-0.2 -1,-0.2 -0.535 59.9 161.1 -89.9 69.8 -7.0 38.7 69.8 105 108 A I H > S+ 0 0 7 -2,-1.7 4,-2.6 1,-0.2 -1,-0.2 0.899 70.7 45.8 -58.5 -47.9 -8.9 37.0 67.0 106 109 A D H > S+ 0 0 110 -3,-0.3 4,-2.4 2,-0.2 -1,-0.2 0.887 111.3 52.9 -65.6 -38.8 -8.4 33.4 68.1 107 110 A K H > S+ 0 0 139 2,-0.2 4,-1.8 1,-0.2 -2,-0.2 0.929 112.3 45.7 -61.4 -45.2 -9.4 34.4 71.7 108 111 A I H X S+ 0 0 2 -4,-2.4 4,-2.6 2,-0.2 5,-0.2 0.920 108.9 55.5 -63.8 -43.6 -12.6 36.0 70.3 109 112 A Y H X S+ 0 0 23 -4,-2.6 4,-2.2 1,-0.2 -2,-0.2 0.937 108.3 48.8 -54.5 -48.1 -13.3 33.0 68.0 110 113 A T H X S+ 0 0 62 -4,-2.4 4,-2.2 1,-0.2 -1,-0.2 0.870 109.5 52.2 -61.8 -36.9 -13.2 30.7 71.1 111 114 A K H X S+ 0 0 70 -4,-1.8 4,-2.1 2,-0.2 -1,-0.2 0.913 109.3 49.3 -65.4 -42.7 -15.5 33.0 73.0 112 115 A L H X S+ 0 0 3 -4,-2.6 4,-1.0 1,-0.2 -2,-0.2 0.872 110.0 52.4 -64.7 -36.8 -18.0 32.9 70.1 113 116 A L H < S+ 0 0 34 -4,-2.2 3,-0.3 -5,-0.2 -2,-0.2 0.910 110.9 46.0 -65.7 -41.8 -17.7 29.1 70.0 114 117 A W H >< S+ 0 0 178 -4,-2.2 3,-1.3 1,-0.2 -2,-0.2 0.849 107.7 57.7 -69.0 -33.2 -18.5 28.8 73.8 115 118 A A H 3< S+ 0 0 49 -4,-2.1 -1,-0.2 1,-0.3 -2,-0.2 0.768 120.9 29.3 -67.1 -24.6 -21.4 31.3 73.4 116 119 A G T >< S+ 0 0 11 -4,-1.0 3,-1.6 -3,-0.3 -1,-0.3 -0.132 79.4 136.5-125.2 37.4 -22.8 28.8 70.8 117 120 A A G X + 0 0 54 -3,-1.3 3,-1.3 1,-0.3 -1,-0.1 0.778 63.6 66.2 -56.2 -31.2 -21.4 25.6 72.2 118 121 A S G 3 S+ 0 0 128 1,-0.2 -1,-0.3 -4,-0.2 -2,-0.1 0.763 99.2 52.1 -64.6 -25.7 -24.7 23.7 71.7 119 122 A V G < S- 0 0 83 -3,-1.6 6,-0.4 1,-0.3 3,-0.3 0.193 108.3-118.2-100.7 16.9 -24.5 24.0 67.9 120 123 A G < - 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