==== Secondary Structure Definition by the program DSSP, updated CMBI version by ElmK / April 1,2000 ==== DATE=13-NOV-2011 . REFERENCE W. KABSCH AND C.SANDER, BIOPOLYMERS 22 (1983) 2577-2637 . HEADER IMMUNE SYSTEM 20-MAY-11 3S4S . COMPND 2 MOLECULE: HLA CLASS II HISTOCOMPATIBILITY ANTIGEN, DR ALPHA . SOURCE 2 ORGANISM_SCIENTIFIC: HOMO SAPIENS; . AUTHOR Y.LI . 1106 12 10 10 0 TOTAL NUMBER OF RESIDUES, NUMBER OF CHAINS, NUMBER OF SS-BRIDGES(TOTAL,INTRACHAIN,INTERCHAIN) . 51273.0 ACCESSIBLE SURFACE OF PROTEIN (ANGSTROM**2) . 776 70.2 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(J) , SAME NUMBER PER 100 RESIDUES . 22 2.0 TOTAL NUMBER OF HYDROGEN BONDS IN PARALLEL BRIDGES, SAME NUMBER PER 100 RESIDUES . 399 36.1 TOTAL NUMBER OF HYDROGEN BONDS IN ANTIPARALLEL BRIDGES, SAME NUMBER PER 100 RESIDUES . 9 0.8 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(I-5), SAME NUMBER PER 100 RESIDUES . 8 0.7 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(I-4), SAME NUMBER PER 100 RESIDUES . 13 1.2 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(I-3), SAME NUMBER PER 100 RESIDUES . 1 0.1 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(I-2), SAME NUMBER PER 100 RESIDUES . 2 0.2 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(I-1), SAME NUMBER PER 100 RESIDUES . 0 0.0 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(I+0), SAME NUMBER PER 100 RESIDUES . 0 0.0 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(I+1), SAME NUMBER PER 100 RESIDUES . 142 12.8 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(I+2), SAME NUMBER PER 100 RESIDUES . 93 8.4 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(I+3), SAME NUMBER PER 100 RESIDUES . 94 8.5 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(I+4), SAME NUMBER PER 100 RESIDUES . 18 1.6 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(I+5), SAME NUMBER PER 100 RESIDUES . 1 2 3 4 5 6 7 8 9 10 11 12 13 14 15 16 17 18 19 20 21 22 23 24 25 26 27 28 29 30 *** HISTOGRAMS OF *** . 0 0 0 1 0 0 0 5 0 0 0 0 1 0 0 0 0 0 0 1 1 0 0 0 0 0 0 0 0 0 RESIDUES PER ALPHA HELIX . 3 1 0 0 1 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 PARALLEL BRIDGES PER LADDER . 9 11 12 9 3 14 1 5 3 2 2 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 ANTIPARALLEL BRIDGES PER LADDER . 5 3 5 2 0 0 2 0 0 2 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 LADDERS PER SHEET . # RESIDUE AA STRUCTURE BP1 BP2 ACC N-H-->O O-->H-N N-H-->O O-->H-N TCO KAPPA ALPHA PHI PSI X-CA Y-CA Z-CA 1 3 A E 0 0 131 0, 0.0 197,-0.2 0, 0.0 3,-0.1 0.000 360.0 360.0 360.0 107.2 -17.0 14.3 -57.6 2 4 A E + 0 0 103 195,-2.8 2,-0.3 1,-0.4 196,-0.2 0.769 360.0 7.3-111.2 -36.2 -15.1 11.0 -58.1 3 5 A H E -A 197 0A 26 194,-1.6 194,-2.9 197,-0.1 -1,-0.4 -0.990 51.6-165.4-148.9 156.8 -12.0 11.3 -55.9 4 6 A V E -A 196 0A 23 -2,-0.3 21,-3.3 192,-0.2 2,-0.5 -0.982 1.2-168.7-146.8 129.2 -10.3 13.5 -53.3 5 7 A I E -AB 195 24A 5 190,-2.2 190,-3.3 -2,-0.4 2,-0.5 -0.969 13.4-164.9-117.2 130.6 -6.8 13.7 -52.0 6 8 A I E -AB 194 23A 24 17,-2.7 17,-2.2 -2,-0.5 2,-0.7 -0.970 16.4-160.3-124.2 129.7 -6.1 15.7 -48.9 7 9 A Q E -AB 193 22A 0 186,-2.3 186,-2.5 -2,-0.5 2,-0.4 -0.927 28.7-179.0 -98.9 117.5 -2.8 16.9 -47.6 8 10 A A E +AB 192 21A 9 13,-3.1 13,-1.9 -2,-0.7 2,-0.3 -0.978 11.2 161.2-130.4 128.4 -3.7 17.7 -43.9 9 11 A E E -AB 191 20A 2 182,-2.2 182,-2.0 -2,-0.4 2,-0.3 -0.879 8.3-179.4-134.7 164.6 -1.4 19.1 -41.2 10 12 A F E -AB 190 19A 8 9,-1.8 9,-1.9 -2,-0.3 2,-0.4 -0.990 17.3-150.0-162.3 158.1 -1.9 20.8 -37.9 11 13 A Y E -AB 189 18A 19 178,-2.3 178,-2.3 -2,-0.3 2,-0.4 -0.999 15.2-164.9-135.7 133.6 -0.1 22.3 -34.9 12 14 A L E >> -AB 188 17A 0 5,-2.5 5,-1.7 -2,-0.4 4,-1.6 -0.975 15.8-124.6-125.2 130.3 -1.5 22.4 -31.4 13 15 A N E 45S+A 187 0A 13 174,-2.2 174,-2.6 -2,-0.4 -2,-0.0 -0.861 82.4 15.0-118.5 154.8 -0.4 24.4 -28.3 14 16 A P T 45S+ 0 0 42 0, 0.0 -1,-0.2 0, 0.0 172,-0.1 -0.991 124.8 61.0 -78.8 3.8 0.5 24.3 -25.6 15 17 A D T 45S- 0 0 20 1,-0.1 -2,-0.3 2,-0.1 3,-0.1 0.717 88.9-141.3 -61.2 -27.3 1.0 20.6 -26.4 16 18 A Q T <5 + 0 0 132 -4,-1.6 2,-0.4 1,-0.2 -3,-0.2 0.912 48.2 149.1 62.6 42.3 3.5 21.2 -29.1 17 19 A S E < -B 12 0A 49 -5,-1.7 -5,-2.5 2,-0.0 2,-0.3 -0.893 22.1-179.7-110.5 137.8 2.0 18.4 -31.2 18 20 A G E -B 11 0A 22 -2,-0.4 2,-0.3 -7,-0.2 16,-0.2 -0.953 6.6-167.5-135.3 154.2 2.1 18.5 -35.0 19 21 A E E -B 10 0A 19 -9,-1.9 -9,-1.8 -2,-0.3 2,-0.4 -0.996 5.7-174.3-144.1 146.9 0.8 16.1 -37.6 20 22 A F E +B 9 0A 0 -2,-0.3 12,-1.8 12,-0.3 2,-0.3 -0.976 22.7 143.1-142.6 125.4 1.2 15.8 -41.4 21 23 A M E -BC 8 31A 0 -13,-1.9 -13,-3.1 -2,-0.4 2,-0.4 -0.978 37.5-136.2-157.3 167.7 -0.6 13.3 -43.6 22 24 A F E -BC 7 30A 0 8,-2.0 7,-2.7 -2,-0.3 8,-1.3 -0.990 22.9-171.5-127.4 134.9 -2.2 12.6 -47.0 23 25 A D E -BC 6 28A 7 -17,-2.2 -17,-2.7 -2,-0.4 2,-0.5 -0.938 13.2-147.4-128.7 153.2 -5.6 10.8 -47.2 24 26 A F E > S-BC 5 27A 2 3,-3.5 3,-2.3 -2,-0.3 -19,-0.2 -0.975 79.4 -26.0-123.0 117.1 -7.4 9.5 -50.3 25 27 A D T 3 S- 0 0 53 -21,-3.3 -1,-0.1 -2,-0.5 -20,-0.1 0.886 131.9 -43.1 47.0 42.6 -11.2 9.5 -50.2 26 28 A G T 3 S+ 0 0 17 -22,-0.3 2,-0.3 1,-0.3 -1,-0.3 0.343 118.5 106.4 90.6 -10.4 -11.1 9.4 -46.4 27 29 A D E < S-C 24 0A 1 -3,-2.3 -3,-3.5 301,-0.1 2,-0.4 -0.827 73.7-111.7-104.9 147.0 -8.4 6.8 -46.0 28 30 A E E -C 23 0A 6 -2,-0.3 -5,-0.3 299,-0.3 14,-0.1 -0.605 25.7-177.2 -78.1 125.4 -4.9 7.5 -44.9 29 31 A I E - 0 0 1 -7,-2.7 13,-2.1 -2,-0.4 14,-0.5 0.891 67.5 -45.6 -83.3 -45.1 -2.2 7.0 -47.5 30 32 A F E -CD 22 41A 0 -8,-1.3 -8,-2.0 11,-0.3 -1,-0.4 -0.973 49.8-153.5-173.9 168.2 0.6 7.9 -45.1 31 33 A H E -CD 21 40A 35 9,-2.2 9,-3.6 -2,-0.3 2,-0.6 -0.888 27.1-106.5-145.1 175.3 1.8 10.3 -42.4 32 34 A V E - D 0 39A 4 -12,-1.8 2,-0.8 -2,-0.3 -12,-0.3 -0.939 19.8-138.1-115.8 116.0 5.1 11.5 -41.1 33 35 A D E >> - D 0 38A 47 5,-2.4 5,-1.7 -2,-0.6 4,-0.8 -0.580 18.4-155.9 -67.1 107.9 6.3 10.3 -37.7 34 36 A M T 45S+ 0 0 47 -2,-0.8 3,-0.3 -16,-0.2 -1,-0.2 0.857 86.9 49.2 -57.3 -41.4 7.7 13.6 -36.4 35 37 A A T 45S+ 0 0 93 1,-0.2 -1,-0.2 2,-0.1 -2,-0.1 0.801 119.9 36.4 -71.7 -30.6 10.1 12.1 -33.9 36 38 A K T 45S- 0 0 145 -3,-0.3 -1,-0.2 2,-0.1 -2,-0.2 0.377 98.6-134.5-101.9 3.7 11.7 9.7 -36.4 37 39 A K T <5 + 0 0 115 -4,-0.8 2,-0.3 -3,-0.3 -3,-0.2 0.915 62.0 141.3 41.6 49.0 11.4 12.2 -39.3 38 40 A E E < -D 33 0A 97 -5,-1.7 -5,-2.4 16,-0.1 2,-0.5 -0.916 58.0-124.5-126.0 148.1 10.0 9.1 -41.1 39 41 A T E -D 32 0A 24 -2,-0.3 2,-0.6 -7,-0.2 -7,-0.2 -0.796 26.6-162.2 -91.5 123.4 7.3 8.5 -43.6 40 42 A V E -D 31 0A 41 -9,-3.6 -9,-2.2 -2,-0.5 2,-0.2 -0.927 6.8-146.9-114.6 116.4 4.8 5.9 -42.5 41 43 A W E -D 30 0A 28 -2,-0.6 -11,-0.3 -11,-0.2 3,-0.2 -0.547 13.3-137.1 -78.0 139.8 2.5 4.2 -45.0 42 44 A R S S+ 0 0 48 -13,-2.1 2,-0.4 1,-0.3 -1,-0.1 0.924 100.9 25.7 -61.3 -43.8 -1.0 3.2 -43.8 43 45 A L S > S- 0 0 30 -14,-0.5 3,-2.0 1,-0.1 4,-0.5 -0.989 85.2-134.9-121.0 126.2 -0.5 -0.1 -45.7 44 46 A E G > S+ 0 0 131 -2,-0.4 3,-2.2 1,-0.3 4,-0.2 0.851 100.0 66.1 -50.5 -44.2 3.0 -1.3 -46.2 45 47 A E G > S+ 0 0 80 1,-0.3 3,-1.6 2,-0.2 4,-0.4 0.773 90.5 68.8 -51.2 -25.8 2.6 -2.2 -49.9 46 48 A F G X> S+ 0 0 0 -3,-2.0 3,-1.3 1,-0.3 4,-0.8 0.863 90.0 62.0 -61.0 -36.5 2.1 1.5 -50.5 47 49 A G G <4 S+ 0 0 34 -3,-2.2 -1,-0.3 -4,-0.5 -2,-0.2 0.364 98.0 56.8 -73.1 6.5 5.7 2.1 -49.7 48 50 A R G <4 S+ 0 0 163 -3,-1.6 -1,-0.2 -4,-0.2 -2,-0.2 0.494 110.5 41.5-111.4 -10.2 6.9 -0.1 -52.6 49 51 A F T <4 S+ 0 0 84 -3,-1.3 2,-0.3 -4,-0.4 -2,-0.2 0.447 125.0 8.1-115.4 -4.5 5.0 1.9 -55.2 50 52 A A < - 0 0 9 -4,-0.8 2,-0.3 216,-0.1 -1,-0.2 -0.978 58.1-154.8-165.3 170.9 5.8 5.4 -54.0 51 53 A S B -h 369 0B 56 317,-1.2 319,-2.3 -2,-0.3 2,-0.4 -0.981 4.9-155.1-153.6 163.0 7.7 7.6 -51.6 52 54 A F - 0 0 16 -2,-0.3 2,-1.0 317,-0.2 -13,-0.2 -0.984 26.7-123.6-144.6 129.6 7.7 10.9 -49.8 53 55 A E > - 0 0 151 317,-0.4 3,-1.3 -2,-0.4 4,-0.4 -0.654 22.1-154.0 -77.1 106.5 10.7 12.8 -48.4 54 56 A A T >> S+ 0 0 11 -2,-1.0 4,-1.4 1,-0.2 3,-0.9 0.693 83.1 73.2 -53.9 -25.7 9.7 13.2 -44.8 55 57 A Q H 3> S+ 0 0 99 1,-0.2 4,-2.0 2,-0.2 -1,-0.2 0.883 86.5 62.8 -61.5 -38.2 11.8 16.4 -44.3 56 58 A G H <> S+ 0 0 17 -3,-1.3 4,-1.7 1,-0.2 -1,-0.2 0.810 102.4 51.6 -56.1 -31.5 9.4 18.5 -46.3 57 59 A A H <> S+ 0 0 0 -3,-0.9 4,-2.7 -4,-0.4 -1,-0.2 0.928 105.2 54.6 -72.1 -44.8 6.8 17.8 -43.7 58 60 A L H X S+ 0 0 39 -4,-1.4 4,-1.8 1,-0.2 -2,-0.2 0.873 109.5 48.6 -56.2 -37.2 9.1 18.8 -40.8 59 61 A A H X S+ 0 0 57 -4,-2.0 4,-1.5 2,-0.2 -1,-0.2 0.891 110.0 51.4 -70.9 -38.4 9.6 22.1 -42.6 60 62 A N H X S+ 0 0 4 -4,-1.7 4,-2.0 -5,-0.2 -2,-0.2 0.923 108.1 52.4 -64.4 -41.0 5.8 22.6 -43.1 61 63 A I H X S+ 0 0 15 -4,-2.7 4,-3.3 1,-0.2 5,-0.2 0.934 104.1 55.8 -60.9 -47.0 5.2 21.9 -39.4 62 64 A A H X S+ 0 0 64 -4,-1.8 4,-1.9 1,-0.3 -1,-0.2 0.886 110.5 45.4 -52.9 -42.4 7.7 24.6 -38.3 63 65 A V H X S+ 0 0 25 -4,-1.5 4,-2.1 2,-0.2 -1,-0.3 0.860 111.4 53.6 -71.1 -32.8 5.9 27.1 -40.3 64 66 A D H X S+ 0 0 0 -4,-2.0 4,-2.4 2,-0.2 -2,-0.2 0.943 106.9 51.2 -65.8 -46.1 2.6 25.8 -38.9 65 67 A K H X S+ 0 0 62 -4,-3.3 4,-1.6 1,-0.2 -2,-0.2 0.939 112.0 47.0 -54.7 -48.8 3.8 26.3 -35.4 66 68 A A H X S+ 0 0 47 -4,-1.9 4,-1.6 1,-0.2 -1,-0.2 0.855 112.1 49.6 -63.1 -37.3 4.8 29.9 -36.1 67 69 A N H X S+ 0 0 3 -4,-2.1 4,-2.9 2,-0.2 -1,-0.2 0.840 103.8 59.5 -72.7 -30.4 1.5 30.6 -37.8 68 70 A L H X S+ 0 0 0 -4,-2.4 4,-2.3 1,-0.2 -2,-0.2 0.909 105.9 49.2 -62.0 -40.9 -0.4 29.2 -34.8 69 71 A E H X S+ 0 0 77 -4,-1.6 4,-2.0 2,-0.2 -1,-0.2 0.908 109.9 52.0 -63.9 -40.8 1.3 31.8 -32.6 70 72 A I H X S+ 0 0 37 -4,-1.6 4,-2.5 1,-0.2 -2,-0.2 0.952 113.6 42.6 -59.9 -51.2 0.4 34.5 -35.1 71 73 A M H X S+ 0 0 0 -4,-2.9 4,-1.7 1,-0.2 -1,-0.2 0.818 108.1 61.5 -65.5 -31.9 -3.3 33.4 -35.1 72 74 A T H <>S+ 0 0 8 -4,-2.3 5,-2.7 -5,-0.2 6,-0.3 0.914 112.3 36.2 -62.2 -42.8 -3.3 33.0 -31.4 73 75 A K H ><5S+ 0 0 109 -4,-2.0 3,-3.3 3,-0.2 -2,-0.2 0.961 112.7 58.1 -73.6 -51.1 -2.4 36.7 -30.8 74 76 A R H 3<5S+ 0 0 58 -4,-2.5 -2,-0.2 1,-0.3 -1,-0.2 0.787 107.1 48.8 -49.2 -33.3 -4.5 38.0 -33.7 75 77 A S T 3<5S- 0 0 11 -4,-1.7 -1,-0.3 -5,-0.2 -2,-0.2 0.450 120.3-109.6 -87.8 0.3 -7.6 36.4 -32.2 76 78 A N T < 5S- 0 0 136 -3,-3.3 -3,-0.2 1,-0.2 -2,-0.2 0.891 72.5 -57.7 71.0 40.7 -6.8 37.9 -28.8 77 79 A Y S S-NO 159 124E 83 3,-3.4 3,-2.0 -2,-0.4 38,-0.2 -0.988 84.3 -17.0-117.9 119.5 -11.3 -6.9 -21.4 122 124 A N T 3 S- 0 0 87 36,-3.2 2,-0.6 -2,-0.5 -1,-0.2 0.959 130.6 -52.7 48.9 55.4 -13.1 -7.9 -18.3 123 125 A G T 3 S+ 0 0 52 35,-0.3 -1,-0.3 -3,-0.2 36,-0.1 0.066 119.0 105.9 72.3 -26.9 -12.1 -4.6 -16.7 124 126 A K E < S-O 121 0E 157 -3,-2.0 -3,-3.4 -2,-0.6 2,-0.2 -0.696 73.0-114.7 -94.1 138.9 -8.4 -5.1 -17.4 125 127 A P E -O 120 0E 87 0, 0.0 2,-0.3 0, 0.0 -5,-0.3 -0.478 29.8-161.3 -71.8 132.7 -6.4 -3.2 -20.1 126 128 A V + 0 0 11 -7,-2.0 -7,-0.5 -2,-0.2 3,-0.0 -0.726 22.4 167.0-110.6 159.8 -5.0 -5.2 -23.0 127 129 A T > + 0 0 93 -2,-0.3 3,-1.5 -9,-0.1 2,-0.6 0.365 31.5 128.0-151.4 -3.2 -2.3 -4.2 -25.5 128 130 A T T 3 S- 0 0 114 1,-0.3 -2,-0.0 21,-0.0 0, 0.0 -0.483 93.9 -13.1 -67.6 112.9 -1.3 -7.4 -27.3 129 131 A G T 3 S+ 0 0 55 -2,-0.6 2,-0.3 1,-0.2 -1,-0.3 0.697 94.9 150.9 71.0 19.3 -1.4 -6.6 -31.0 130 132 A V < + 0 0 49 -3,-1.5 2,-0.3 19,-0.2 -1,-0.2 -0.670 17.7 176.2 -86.5 138.2 -3.4 -3.3 -30.6 131 133 A S E -L 148 0D 66 17,-1.0 17,-3.5 -2,-0.3 2,-0.3 -0.796 8.3-168.6-131.1 173.6 -2.9 -0.6 -33.2 132 134 A E E -L 147 0D 116 15,-0.3 15,-0.3 -2,-0.3 2,-0.2 -0.984 17.8-121.5-158.4 162.6 -4.3 2.8 -33.9 133 135 A T - 0 0 6 13,-2.8 2,-0.1 -2,-0.3 3,-0.1 -0.509 30.5-109.9-101.2 174.6 -4.4 5.5 -36.6 134 136 A V - 0 0 25 -2,-0.2 2,-0.6 1,-0.1 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