==== Secondary Structure Definition by the program DSSP, updated CMBI version by ElmK / April 1,2000 ==== DATE=20-JUL-2011 . REFERENCE W. KABSCH AND C.SANDER, BIOPOLYMERS 22 (1983) 2577-2637 . HEADER TRANSFERASE 28-SEP-07 2VD3 . COMPND 2 MOLECULE: ATP PHOSPHORIBOSYLTRANSFERASE; . SOURCE 2 ORGANISM_SCIENTIFIC: METHANOBACTERIUM THERMOAUTOTROPHICUM; . AUTHOR B.LOHKAMP,T.SCHWEIKERT,A.J.LAPTHORN . 579 5 0 0 0 TOTAL NUMBER OF RESIDUES, NUMBER OF CHAINS, NUMBER OF SS-BRIDGES(TOTAL,INTRACHAIN,INTERCHAIN) . 29846.0 ACCESSIBLE SURFACE OF PROTEIN (ANGSTROM**2) . 393 67.9 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(J) , SAME NUMBER PER 100 RESIDUES . 36 6.2 TOTAL NUMBER OF HYDROGEN BONDS IN PARALLEL BRIDGES, SAME NUMBER PER 100 RESIDUES . 116 20.0 TOTAL NUMBER OF HYDROGEN BONDS IN ANTIPARALLEL BRIDGES, SAME NUMBER PER 100 RESIDUES . 2 0.3 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(I-5), SAME NUMBER PER 100 RESIDUES . 0 0.0 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(I-4), SAME NUMBER PER 100 RESIDUES . 0 0.0 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(I-3), SAME NUMBER PER 100 RESIDUES . 1 0.2 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(I-2), SAME NUMBER PER 100 RESIDUES . 0 0.0 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(I-1), SAME NUMBER PER 100 RESIDUES . 0 0.0 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(I+0), SAME NUMBER PER 100 RESIDUES . 0 0.0 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(I+1), SAME NUMBER PER 100 RESIDUES . 39 6.7 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(I+2), SAME NUMBER PER 100 RESIDUES . 44 7.6 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(I+3), SAME NUMBER PER 100 RESIDUES . 127 21.9 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(I+4), SAME NUMBER PER 100 RESIDUES . 14 2.4 TOTAL NUMBER OF HYDROGEN BONDS OF TYPE O(I)-->H-N(I+5), SAME NUMBER PER 100 RESIDUES . 1 2 3 4 5 6 7 8 9 10 11 12 13 14 15 16 17 18 19 20 21 22 23 24 25 26 27 28 29 30 *** HISTOGRAMS OF *** . 0 0 0 1 2 2 6 0 1 3 1 0 1 0 1 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 RESIDUES PER ALPHA HELIX . 0 4 0 2 0 2 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 PARALLEL BRIDGES PER LADDER . 0 2 2 4 6 0 0 2 2 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 ANTIPARALLEL BRIDGES PER LADDER . 0 0 2 2 0 2 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 LADDERS PER SHEET . # RESIDUE AA STRUCTURE BP1 BP2 ACC N-H-->O O-->H-N N-H-->O O-->H-N TCO KAPPA ALPHA PHI PSI X-CA Y-CA Z-CA 1 -1 A A 0 0 163 0, 0.0 0, 0.0 0, 0.0 0, 0.0 0.000 360.0 360.0 360.0 -65.7 62.1 61.4 88.7 2 0 A V - 0 0 109 0, 0.0 2,-0.3 0, 0.0 0, 0.0 -0.990 360.0-114.8-139.4 138.6 60.3 64.8 88.8 3 1 A P - 0 0 53 0, 0.0 2,-0.3 0, 0.0 196,-0.1 -0.572 18.3-157.0 -90.6 138.3 57.6 66.3 86.6 4 2 A K - 0 0 142 -2,-0.3 2,-0.4 43,-0.0 43,-0.2 -0.783 9.7-152.4 -98.4 146.9 57.8 69.3 84.2 5 3 A I E -a 47 0A 2 41,-3.3 43,-2.8 -2,-0.3 2,-0.5 -0.939 3.4-152.3-119.9 149.9 54.8 71.3 83.0 6 4 A R E -a 48 0A 60 -2,-0.4 62,-1.8 41,-0.2 61,-1.6 -0.979 10.2-160.8-118.2 123.3 54.3 73.3 79.9 7 5 A I E -ab 49 68A 0 41,-2.7 43,-2.7 -2,-0.5 2,-0.5 -0.929 13.8-136.9-107.7 128.7 51.9 76.2 79.9 8 6 A A E -ab 50 69A 0 60,-3.1 62,-2.1 -2,-0.5 43,-0.2 -0.711 26.3-177.7 -84.3 121.0 50.4 77.6 76.7 9 7 A V E -a 51 0A 0 41,-2.5 43,-2.0 -2,-0.5 62,-0.1 -0.970 33.2-101.8-125.1 133.2 50.5 81.4 76.6 10 8 A P E -a 52 0A 0 0, 0.0 43,-0.2 0, 0.0 10,-0.1 -0.264 24.6-158.7 -55.7 141.1 49.1 83.6 73.8 11 9 A S S S+ 0 0 52 41,-1.0 2,-0.3 1,-0.1 42,-0.1 0.741 70.8 16.7 -94.3 -32.3 51.8 85.0 71.6 12 10 A K S S+ 0 0 133 40,-0.2 42,-0.3 4,-0.1 3,-0.2 -0.875 94.2 43.5-135.9 169.4 49.9 88.0 70.1 13 11 A G S >> S- 0 0 64 -2,-0.3 3,-0.8 1,-0.2 4,-0.6 -0.204 98.0 -47.6 87.5-177.1 46.7 90.0 70.9 14 12 A R T 34 S+ 0 0 136 1,-0.2 -1,-0.2 2,-0.1 78,-0.1 0.772 128.9 55.2 -66.4 -32.6 45.2 91.5 74.0 15 13 A I T 3> S+ 0 0 17 -3,-0.2 4,-1.5 1,-0.1 -1,-0.2 0.765 94.0 74.5 -70.0 -27.7 45.5 88.4 76.2 16 14 A S H <> S+ 0 0 11 -3,-0.8 4,-1.8 -4,-0.2 3,-0.4 0.935 94.0 39.4 -60.6 -66.8 49.3 88.0 75.7 17 15 A E H X S+ 0 0 137 -4,-0.6 4,-2.7 1,-0.2 -1,-0.2 0.862 114.9 51.1 -54.8 -47.3 51.1 90.7 77.7 18 16 A P H > S+ 0 0 32 0, 0.0 4,-2.4 0, 0.0 -1,-0.2 0.855 109.0 53.4 -61.9 -33.1 48.8 90.5 80.8 19 17 A A H X S+ 0 0 0 -4,-1.5 4,-2.0 -3,-0.4 -2,-0.2 0.888 110.1 47.1 -63.9 -41.4 49.3 86.7 80.9 20 18 A I H X S+ 0 0 49 -4,-1.8 4,-2.2 2,-0.2 10,-0.2 0.931 113.4 48.7 -65.1 -44.6 53.1 87.2 80.9 21 19 A R H X S+ 0 0 133 -4,-2.7 4,-2.5 2,-0.2 -2,-0.2 0.860 109.4 52.7 -62.2 -39.4 52.8 89.9 83.6 22 20 A L H X S+ 0 0 14 -4,-2.4 4,-1.5 2,-0.2 6,-0.2 0.932 109.6 47.1 -64.0 -48.0 50.6 87.6 85.8 23 21 A L H <>S+ 0 0 2 -4,-2.0 5,-3.1 2,-0.2 4,-0.3 0.917 110.9 53.9 -58.4 -45.3 53.1 84.7 85.6 24 22 A E H ><5S+ 0 0 106 -4,-2.2 3,-1.5 4,-0.3 5,-0.5 0.944 110.6 45.3 -50.1 -52.8 55.8 87.2 86.5 25 23 A N H 3<5S+ 0 0 59 -4,-2.5 -1,-0.2 1,-0.3 -2,-0.2 0.732 107.6 59.3 -66.5 -22.4 53.8 88.3 89.6 26 24 A A T 3<5S- 0 0 7 -4,-1.5 -1,-0.3 -3,-0.2 -2,-0.2 0.398 123.5-102.7 -84.9 -0.3 53.2 84.6 90.5 27 25 A G T < 5S+ 0 0 35 -3,-1.5 -3,-0.2 -4,-0.3 -2,-0.1 0.543 102.1 100.1 87.2 11.0 56.9 84.0 90.7 28 26 A V < + 0 0 8 -5,-3.1 -4,-0.3 -6,-0.2 -3,-0.1 -0.296 48.2 159.2-124.5 44.2 57.0 82.2 87.3 29 27 A G - 0 0 0 -5,-0.5 14,-1.9 -6,-0.3 15,-0.2 -0.272 53.4 -84.0 -65.7 155.3 58.3 85.0 85.0 30 28 A L E -C 42 0A 44 12,-0.2 2,-0.6 -10,-0.2 12,-0.3 -0.389 47.0-147.9 -61.6 133.7 59.9 84.1 81.7 31 29 A K E -C 41 0A 78 10,-3.2 10,-0.6 1,-0.2 -1,-0.1 -0.930 62.8 -24.5-118.4 114.7 63.5 83.2 82.1 32 30 A D S S- 0 0 125 -2,-0.6 2,-0.7 1,-0.2 -1,-0.2 0.837 76.8-169.4 62.5 38.8 66.2 83.9 79.5 33 31 A T - 0 0 48 8,-0.1 2,-0.2 -3,-0.1 -1,-0.2 -0.492 6.5-154.4 -69.6 105.7 63.7 84.0 76.6 34 32 A V > - 0 0 108 -2,-0.7 3,-0.8 5,-0.2 2,-0.2 -0.560 23.3-111.3 -74.4 141.9 65.6 84.0 73.3 35 33 A N T 3 S+ 0 0 142 1,-0.2 3,-0.1 -2,-0.2 -1,-0.1 -0.511 102.7 37.7 -75.4 149.0 63.7 85.6 70.4 36 34 A R T 3 S+ 0 0 268 1,-0.3 2,-0.5 -2,-0.2 -1,-0.2 0.511 91.5 130.2 81.2 10.3 62.6 83.2 67.6 37 35 A K < - 0 0 38 -3,-0.8 -1,-0.3 1,-0.1 14,-0.1 -0.839 46.6-168.5-101.7 130.6 61.8 80.7 70.3 38 36 A L S S+ 0 0 73 -2,-0.5 13,-2.7 -3,-0.1 2,-0.3 0.671 81.3 46.7 -85.4 -22.1 58.5 78.9 70.5 39 37 A F E - D 0 50A 53 11,-0.2 2,-0.3 -5,-0.1 11,-0.2 -0.867 67.8-174.6-117.2 155.1 59.4 77.7 74.0 40 38 A S E - D 0 49A 10 9,-2.0 9,-3.0 -2,-0.3 -8,-0.1 -0.990 22.5-124.6-147.6 140.6 60.8 79.5 77.0 41 39 A K E -CD 31 48A 128 -10,-0.6 -10,-3.2 -2,-0.3 2,-0.2 -0.349 27.8-150.0 -70.4 162.3 62.1 78.6 80.4 42 40 A T E -C 30 0A 4 5,-0.9 -12,-0.2 2,-0.5 -13,-0.1 -0.713 35.5 -90.3-123.6 179.6 60.6 80.4 83.5 43 41 A Q S S+ 0 0 108 -14,-1.9 -15,-0.1 -2,-0.2 -13,-0.1 0.690 112.4 74.4 -66.2 -18.1 61.9 81.3 87.0 44 42 A H S > S- 0 0 29 -15,-0.2 3,-1.9 3,-0.2 -2,-0.5 -0.865 81.2-141.9 -97.6 124.2 60.6 77.9 88.1 45 43 A P T 3 S+ 0 0 115 0, 0.0 -1,-0.1 0, 0.0 -2,-0.0 0.634 100.2 48.6 -61.4 -14.4 62.8 75.0 86.9 46 44 A Q T 3 S+ 0 0 51 -42,-0.1 -41,-3.3 1,-0.1 2,-0.5 0.359 98.7 79.2-106.1 1.9 59.8 72.7 86.3 47 45 A I E < +a 5 0A 5 -3,-1.9 -5,-0.9 -43,-0.2 2,-0.3 -0.949 51.9 175.4-118.8 121.9 57.7 75.2 84.4 48 46 A E E -aD 6 41A 58 -43,-2.8 -41,-2.7 -2,-0.5 2,-0.5 -0.836 21.3-133.9-115.1 161.0 58.1 76.1 80.7 49 47 A V E -aD 7 40A 0 -9,-3.0 -9,-2.0 -2,-0.3 2,-0.6 -0.946 8.6-160.2-116.9 129.9 56.1 78.3 78.4 50 48 A M E -aD 8 39A 0 -43,-2.7 -41,-2.5 -2,-0.5 2,-0.7 -0.963 17.7-138.1-105.5 120.2 54.9 77.5 74.9 51 49 A F E +a 9 0A 38 -13,-2.7 2,-0.3 -2,-0.6 -43,-0.1 -0.671 37.8 162.6 -77.4 113.8 53.9 80.5 72.8 52 50 A S E -a 10 0A 5 -43,-2.0 -41,-1.0 -2,-0.7 -40,-0.2 -0.928 45.0 -97.8-131.0 149.9 50.8 79.5 71.0 53 51 A R >> - 0 0 108 -2,-0.3 3,-1.8 -43,-0.2 4,-0.9 -0.502 40.0-118.2 -66.4 140.6 48.1 81.5 69.2 54 52 A A T 34 S+ 0 0 21 -42,-0.3 3,-0.3 1,-0.3 18,-0.1 0.753 110.2 53.7 -56.3 -34.2 45.1 82.0 71.4 55 53 A A T 34 S+ 0 0 35 1,-0.2 4,-0.3 2,-0.1 -1,-0.3 0.672 108.7 51.2 -76.9 -16.0 42.5 80.1 69.3 56 54 A D T <> S+ 0 0 8 -3,-1.8 4,-2.4 1,-0.1 5,-0.2 0.617 88.4 82.7 -93.5 -17.7 44.8 77.0 69.3 57 55 A I H X S+ 0 0 0 -4,-0.9 4,-2.2 -3,-0.3 5,-0.3 0.881 83.7 57.3 -59.9 -42.8 45.3 76.9 73.1 58 56 A P H > S+ 0 0 0 0, 0.0 4,-2.5 0, 0.0 -1,-0.2 0.965 113.5 41.1 -51.9 -50.2 42.0 75.1 73.9 59 57 A E H > S+ 0 0 32 -4,-0.3 4,-2.7 2,-0.2 6,-0.3 0.899 111.3 54.4 -63.6 -45.3 43.1 72.2 71.6 60 58 A F H X>S+ 0 0 15 -4,-2.4 6,-1.9 1,-0.2 5,-1.7 0.876 115.6 41.6 -55.6 -38.7 46.8 72.1 72.6 61 59 A V H <5S+ 0 0 0 -4,-2.2 -2,-0.2 -5,-0.2 -1,-0.2 0.871 112.8 52.5 -77.0 -38.6 45.5 71.7 76.2 62 60 A A H <5S+ 0 0 21 -4,-2.5 -2,-0.2 -5,-0.3 128,-0.2 0.890 115.7 41.5 -62.5 -37.7 42.8 69.2 75.3 63 61 A D H <5S- 0 0 64 -4,-2.7 -1,-0.2 -5,-0.1 -2,-0.2 0.715 114.7-111.7 -86.9 -24.8 45.3 67.0 73.5 64 62 A G T <5S+ 0 0 37 -4,-1.2 -3,-0.2 2,-0.4 3,-0.1 0.542 82.4 122.2 103.1 10.3 48.1 67.3 76.0 65 63 A A S - E 0 184A 2 -2,-0.3 4,-2.0 112,-0.2 3,-0.3 -0.520 30.8-132.5 -83.1 144.1 39.7 83.3 76.4 73 71 A Y H > S+ 0 0 77 110,-2.2 4,-3.0 1,-0.2 5,-0.2 0.851 106.7 60.0 -60.7 -38.6 36.5 81.6 77.4 74 72 A D H > S+ 0 0 11 2,-0.2 4,-2.9 1,-0.2 -1,-0.2 0.906 107.1 45.7 -54.9 -43.5 34.9 82.4 74.0 75 73 A L H > S+ 0 0 34 -3,-0.3 4,-2.4 2,-0.2 5,-0.2 0.926 112.0 51.0 -69.4 -43.9 37.7 80.5 72.3 76 74 A I H X>S+ 0 0 9 -4,-2.0 4,-1.0 1,-0.2 5,-0.9 0.919 115.6 42.7 -56.3 -46.6 37.4 77.5 74.7 77 75 A V H <5S+ 0 0 53 -4,-3.0 3,-0.3 2,-0.2 -2,-0.2 0.925 111.1 54.3 -68.9 -43.4 33.7 77.4 74.1 78 76 A E H <5S+ 0 0 49 -4,-2.9 -2,-0.2 1,-0.3 -1,-0.2 0.906 113.7 41.5 -58.0 -46.6 33.9 77.9 70.3 79 77 A R H <5S- 0 0 66 -4,-2.4 -1,-0.3 -5,-0.1 -2,-0.2 0.710 96.7-143.3 -69.5 -23.3 36.2 75.0 69.9 80 78 A G T <5 + 0 0 62 -4,-1.0 -3,-0.2 -3,-0.3 -4,-0.1 0.771 35.6 179.4 59.7 27.8 34.2 72.9 72.4 81 79 A S < - 0 0 16 -5,-0.9 2,-1.2 1,-0.1 -1,-0.2 -0.209 35.3-114.7 -64.1 148.3 37.6 71.5 73.5 82 80 A D + 0 0 120 -3,-0.1 2,-0.3 -23,-0.1 108,-0.3 -0.706 63.1 134.4 -90.2 94.7 37.7 69.0 76.3 83 81 A V - 0 0 21 -2,-1.2 2,-0.5 106,-0.1 106,-0.2 -0.944 55.3-117.3-139.3 154.5 39.6 70.9 79.1 84 82 A E E -F 188 0A 61 104,-3.1 104,-2.4 -2,-0.3 2,-0.6 -0.862 25.3-140.3 -92.2 124.9 39.4 71.6 82.8 85 83 A I E +F 187 0A 81 -2,-0.5 102,-0.3 102,-0.2 3,-0.1 -0.783 24.3 176.0 -83.5 120.3 38.9 75.3 83.8 86 84 A L E - 0 0 18 100,-3.2 2,-0.3 -2,-0.6 101,-0.2 0.861 57.5 -3.6 -92.4 -43.9 41.0 76.0 86.9 87 85 A E E -F 186 0A 28 99,-1.3 99,-2.7 120,-0.0 2,-0.4 -0.995 57.4-123.5-150.6 146.3 40.7 79.7 87.5 88 86 A D E -F 185 0A 42 -2,-0.3 97,-0.2 97,-0.2 123,-0.0 -0.826 23.4-153.7 -86.1 136.4 39.1 82.8 86.1 89 87 A L - 0 0 8 95,-3.3 96,-0.1 -2,-0.4 -1,-0.1 0.418 21.9-132.0 -99.4 0.5 41.9 85.3 85.6 90 88 A K S S+ 0 0 80 94,-0.4 2,-0.3 1,-0.2 3,-0.1 0.616 72.2 87.6 64.0 20.2 39.8 88.5 85.9 91 89 A Y + 0 0 41 93,-0.2 93,-2.0 1,-0.1 -2,-0.2 -0.889 64.4 43.4-136.3 170.5 41.1 90.3 82.8 92 90 A G S S+ 0 0 16 -2,-0.3 2,-0.3 1,-0.2 -1,-0.1 0.873 71.2 175.1 63.7 38.8 40.2 90.4 79.0 93 91 A R + 0 0 129 90,-0.1 2,-0.3 -3,-0.1 90,-0.2 -0.583 5.6 153.5 -82.6 139.5 36.5 90.6 79.7 94 92 A A E -G 182 0B 8 88,-1.8 88,-2.6 -2,-0.3 2,-0.5 -0.975 35.8-127.1-164.6 148.6 34.3 91.0 76.7 95 93 A S E -GH 181 162B 28 67,-2.2 67,-1.8 -2,-0.3 2,-1.0 -0.922 9.8-144.7-108.3 128.0 30.8 90.3 75.5 96 94 A L E -GH 180 161B 1 84,-2.6 83,-2.6 -2,-0.5 84,-1.5 -0.839 37.9-176.5 -84.4 105.4 29.9 88.4 72.4 97 95 A V E -GH 178 160B 6 63,-3.0 63,-1.7 -2,-1.0 2,-0.5 -0.673 30.5-127.1-104.3 157.0 26.7 90.4 71.5 98 96 A L E -GH 177 159B 0 79,-3.1 78,-3.1 -2,-0.2 79,-1.3 -0.930 38.3-175.3 -99.0 125.9 24.1 90.1 68.8 99 97 A A E -GH 175 158B 0 59,-2.7 59,-2.3 -2,-0.5 76,-0.2 -0.912 14.3-163.8-129.1 150.8 23.7 93.4 67.0 100 98 A A E -G 174 0B 0 74,-2.2 74,-2.8 -2,-0.3 57,-0.1 -0.941 42.7 -83.2-129.2 148.2 21.5 94.9 64.3 101 99 A P E > -G 173 0B 40 0, 0.0 3,-3.0 0, 0.0 72,-0.3 -0.365 50.1-122.8 -42.2 121.0 21.6 98.1 62.1 102 100 A E T 3 S+ 0 0 67 70,-2.3 71,-0.1 1,-0.3 70,-0.1 0.748 108.9 55.4 -52.5 -31.5 20.1 100.6 64.5 103 101 A D T 3 S+ 0 0 153 69,-0.3 -1,-0.3 71,-0.0 70,-0.1 0.389 87.8 119.4 -80.4 3.8 17.3 101.6 62.1 104 102 A S S < S- 0 0 35 -3,-3.0 -4,-0.1 2,-0.2 52,-0.0 -0.242 71.2-129.5 -73.7 156.8 16.1 98.0 61.9 105 103 A T S S+ 0 0 120 6,-0.1 2,-0.3 2,-0.0 -1,-0.1 0.927 87.2 93.7 -60.6 -48.3 12.7 96.5 62.9 106 104 A I + 0 0 13 1,-0.2 -2,-0.2 5,-0.1 -6,-0.1 -0.291 39.2 156.3 -56.1 109.6 14.6 93.8 64.9 107 105 A R + 0 0 162 1,-0.3 -1,-0.2 -2,-0.3 -7,-0.1 0.529 68.3 12.2-110.6 -14.1 14.8 95.1 68.5 108 106 A G S > S- 0 0 5 69,-0.0 3,-2.4 1,-0.0 -1,-0.3 -0.970 88.0 -92.8-156.2 162.6 15.2 91.7 70.3 109 107 A P G > S+ 0 0 20 0, 0.0 3,-1.1 0, 0.0 68,-0.1 0.691 116.4 70.9 -53.7 -20.2 15.9 88.0 69.3 110 108 A E G 3 S+ 0 0 152 1,-0.2 -4,-0.0 25,-0.0 0, 0.0 0.723 92.7 57.7 -67.4 -21.5 12.1 87.4 69.1 111 109 A D G < S+ 0 0 65 -3,-2.4 -1,-0.2 2,-0.1 -5,-0.1 0.477 73.9 118.5 -91.3 -6.6 11.9 89.5 65.9 112 110 A I < - 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